Talacasto Formation

Talacasto Formation
Talacasto Formation
Stratigraphic range: Devonian
Type	Geological formation
Unit of	Gualilán Group
Thickness	300–1,000 m (980–3,280 ft)
Location
Region	San Juan Province; La Rioja Province;
Country	Argentina
Extent	Precordillera Basin
Type section
Named by	Padula et al
Year defined	1967

The Talacasto Formation is a geological formation in Argentina, exposed primarily across San Juan Province, with an additional outcrop in La Rioja Province. It preserves fossils dating back to the Devonian period.

Description

The Talacasto Formation is exposed in the Precordillera Basin of Argentina, with outcrops most widespread in San Juan Province, though an isolated exposure also exists at Sierra de las Minitas in La Rioja Province.¹² Along with the Punta Negra Formation, it makes up the Gualilán Group, which represent Devonian-aged deposits in the Argentine Precordillera.³ The thickness of the Talacasto Formation varies across its extent; at the type locality of Quebrada de Talacasto in the south it is 300 m (980 ft) thick, while at the Loma de los Piojos section near the Jáchal River in the north it exceeds 1,000 m (3,300 ft) in thickness.¹

Fossil content

Color key

Taxon	Reclassified taxon	Taxon falsely reported as present	Dubious taxon or junior synonym	Ichnotaxon	Ootaxon	Morphotaxon

Notes
Uncertain or tentative taxa are in small text; ~~crossed out~~ taxa are discredited.

Bivalves

Bivalves reported from the Talacasto Formation
Genus	Species	Presence	Material	Notes	Images
Actinopteria⁴	A. eschwegei	Loma de los Piojos	About 20 specimens	A pterineid
Actinopteria⁴	A. modesta	Loma de los Piojos	Several specimens	A pterineid
Anthracoleda⁴	A. (Pseudoleda) minuta	Cerro del Fuerte Loma de los Piojos	Several specimens	A nuculanid
Deceptrix⁴	D. (Devonodeceptrix) jachalensis	Loma de los Piojos	Several specimens	A cardiolariid
Glossites⁴	G. cf. G. depressus	Loma de los Piojos	2 specimens	A modiomorphid
Grammysioidea⁴	G. capricornus	Loma de los Piojos	5 specimens	A grammysiid
Nuculites⁴	N. argentinum	Cerro del Fuerte Loma de los Piojos	Numerous specimens	A cucullellid
Nuculoidea⁴	N.? sp.	Cerro del Fuerte	4 specimens	A nuculid
Palaeoneilo⁴	P. cf. P. forbesi	Loma de los Piojos	5 internal moulds	A malletiid
Pholadella⁴	P. radiata	Loma de los Piojos	7 specimens	A sanguinolitid

Brachiopods

Brachiopods reported from the Talacasto Formation
Genus	Species	Presence	Material	Notes	Images
Anoplia⁵⁶	A. azucenae	Portezuelo del Tambolar	Over 70 specimens	An anopliid
	A. sp. A	Río Talacasto	6 specimens	An anopliid
	A. sp.	Los Algarrobos section		An anopliid
Athyris⁶	A. sp.	Los Algarrobos section		An athyridid
Australostrophia⁵	A. penoensis	Loma de los Piojos Cerro del Fuerte Agua del Peñón	Numerous specimens	A strophochonetid
Boucotia⁷	B. argentina	Loma de los Piojos Portezuelo del Tambolar	3 isolated valves	A notanopliid
Chonostrophia⁵⁸	C. elenae	Loma de los Piojos	Several specimens	A chonostrophiid
Chonostrophia⁵⁸	C. reversa var. andina	Quebrada de Talacasto	Several specimens	Tentatively reclassified as a species of Sanjuanetes⁹
Eoschucertella¹⁰	E. delicata	Quebrada Zerina	Numerous specimens	An areostrophiid
Iridistrophia¹⁰	I. aliciae	Cerro del Fuerte	Numerous specimens	A chilidiopsid
	I. dominans	Loma de los Piojos	Numerous specimens	A chilidiopsid
	I. magna	Cordón de Agua de Peñón	Numerous specimens	A chilidiopsid
	I. prima	Quebrada La Cortadera	Numerous specimens	A chilidiopsid
Kentronetes⁵	K. giolittii	Quebrada Las Loras	6 valves	A strophochonetid
	K. ortegae	Loma de los Piojos	Numerous specimens	A strophochonetid
	K. vallensis	Loma de los Piojos	31 specimens	A strophochonetid
	K. variabilis	Portezuelo del Tambolar Poblete Sur Quebrada del río Talacasto Quebrada del Chañar (Cerro La Chilca) Loma de los Piojos	32 valves	A strophochonetid
Lingula⁶	L. sp.	Los Algarrobos section		A lingulid
Lomaella⁵	L. primoris	Loma de los Piojos	9 specimens	An eodevonariid
Lomaella⁵	L. sanjuanina	Loma de los Piojos	Several specimens	An eodevonariid
Meristella⁶	M. sp.	Los Algarrobos section		A meristellid
Metaplasia⁶	M. baldisi	Los Algarrobos section		An ambocoeliid
Pleurochonetes⁵	?P. sp.	Loma de los Piojos	32 specimens	A chonetid
Plicanoplia⁵	P. sp.	Loma de los Piojos	5 specimens	An anopliid
Sanjuanetes⁵⁶⁸	?S. andina	Quebrada de Talacasto Los Algarrobos section	Several specimens	A strophochonetid, originally reported as Chonostrophia reversa var. andina⁹

Cephalopods

Cephalopods reported from the Talacasto Formation
Genus	Species	Presence	Material	Notes	Images
Bactrites¹	B. gracilis	Quebrada de Talacasto Below Keidel's Bed	18 specimens	A bactritid
Bactrites¹	B. sp.	Quebrada de Talacasto Sierra de las Minitas	5 specimens	A bactritid
Devonobactrites¹	D.? sp.	Loma de los Piojos section	2 specimens	A bactritid

Corals

Corals reported from the Talacasto Formation
Genus	Species	Presence	Material	Notes	Images
Enterolasma⁶	E. sp.	Cerro del Fuerte section	Fragments & complete specimens	A horn coral
Favosites⁶	F.? sp.	Cerro del Fuerte section	1 colony	A tabulate coral
Parastriatopora⁶¹¹	P. cf. gigantea	Tambolar section	6 small branch fragments	A tabulate coral
	P. sanjuanina	Los Algarrobos section		A tabulate coral
	P. sp.	Los Algarrobos section		A tabulate coral

Echinoderms

Echinoderms reported from the Talacasto Formation
Genus	Species	Presence	Material	Notes	Images
Argentinaster¹²	A. bodenbenderi	Jachal	Several specimens	An ophiurinid brittle star
Crinoidea indet.⁶	Indeterminate	Los Algarrobos section		A crinoid
Furcaster¹²	F. separatus	Jachal	Multiple specimens	A protasterid brittle star
Marginix¹²	M. yachalensis	Jachal	Several specimens	An encrinasterid brittle star
Promopalaeaster¹²	P.? quadriserialis	Jachal	Several specimens	A valvatid starfish

Hyoliths

Hyoliths reported from the Talacasto Formation
Genus	Species	Presence	Material	Notes	Images
Hyolithida indet.⁶	Indeterminate	Los Algarrobos section		A hyolithid

Ostracods

Ostracods reported from the Talacasto Formation
Genus	Species	Presence	Material	Notes	Images
Aechmina¹³	A. sp.	Talacasto Creek	6 specimens	A member of Podocopa
Amphizona¹³	A.? argentinensis	Portezuelo del Tambolar Talacasto Creek Poblete Sur Las Tunas	23 specimens	A member of Podocopa
Bollia¹³	B. talacastensis	Talacasto Creek Poblete Sur	20 specimens	A member of Podocopa
Keslingiella¹³	K. teresae	Los Algarrobos Talacasto Creek Poblete Sur La Chilcas Las Tunas	Several specimens	A member of Podocopida
Lapazites¹³	L. aff. L. trinodis	Portezuelo del Tambolar La Cortadera Poblete Sur La Chilca	12 specimens	A member of Podocopa
Petrisigmoopsis¹³	P.? rotundum	Talacasto Creek La Chilca	Several specimens	A member of Podocopa
Pircawayra¹³¹⁴	P. antiqua	Portezuelo del Tambolar La Cortadera Creek La Chilca Las Aguaditas Creek	5 specimens	A member of Palaeocopida
Pircawayra¹³¹⁴	P. peregrina	Loma de los Piojos	Several specimens	A member of Palaeocopida
Suinella¹³	S. huarpesi	Portezuelo del Tambolar Talacasto Creek Poblete Sur	Several specimens	A member of Podocopa
Thlipsurella¹³	T. aff. T. putea	Los Algarrobos Talacasto Creek Poblete Sur Las Tunas	33 specimens	A member of Podocopida
Ulrichia¹³	U. sp.	Talacasto Creek Poblete Sur La Chilca	7 specimens	A member of Podocopa

Sponges

Sponges reported from the Talacasto Formation
Genus	Species	Presence	Material	Notes	Images
Talacastospongia¹⁵	T. minima	Talacasto section Los Algarrobos Las Aguaditas	13 specimens	A glass sponge

Trilobites

Trilobites reported from the Talacasto Formation
Genus	Species	Presence	Material	Notes	Images
Aguaditaspis¹⁶	A. mediaspina	Quebrada de las Aguaditas Sierra de las Minitas	Several specimens	A dalmanitid
Bainella¹⁷	B. cooperi	Cerro La Chilca	6 specimens	A calmoniid
Bainella¹⁷	B. sanjuanina	Cerro La Chilca	8 specimens	A calmoniid
Calmonia¹⁷	C. nov. sp. A	Loma de los Piojos	An internal mould (CORD PZ 8613)	A calmoniid
Dalmanitoides¹⁸	D. boehmi	Loma de los Piojos	Several specimens	A dalmanitid
Dalmanitoides¹⁸	D. drevermanni	Loma de los Piojos Cerro La Chilca	Several specimens	A dalmanitid
Dalmanitidae indet.⁶¹⁶	Gen. et. sp. nov.	Quebrada de las Aguaditas	One specimen (CEGH-UNC 27422)	An unnamed dalmanitid
Dalmanitidae indet.⁶¹⁶	Indeterminate	Los Algarrobos section		A dalmanitid
Dalmanitinae gen. et. sp. indet.²	Indeterminate	Sierra de las Minitas	Fragmentary pygidium (PULR-132 A-B)	A dalmanitid
Deltacephalaspis¹⁷	D. sp.	Cerro La Chilca	2 specimens	A calmoniid
Echidnops²¹⁹	E. taphomimus	Quebrada de Las Aguaditas Sierra de Los Blaquitos Sierra de las Minitas	12 specimens	A phacopid, originally reported as an unnamed species of Paciphacops
Ivanites¹⁶	I. leonorae	Quebrada de Las Aguaditas Quebrada de Talacasto Loma de los Piojos	Several specimens	A dalmanitid
Maurotarion⁶²⁰²¹	M. (Malvinotarion) gaucho	Loma de los Piojos	10 specimens	An aulacopleurid
	M. (Malvinotarion) haudei	Loma de los Piojos	2 specimens	An aulacopleurid
	M. (Malvinotarion) talacastoense	La Cantera point	5 specimens	An aulacopleurid
	M. (Malvinotarion) sp.	Portezuelo del Tambolar	One specimen (CEGH-UNC 7.086)	An aulacopleurid
	M. (Malvinotarion) sp. B	Cerro La Chilca	One specimen	An aulacopleurid
	M. megacephalum	Loma de los Piojos	23 specimens	An aulacopleurid
	M. sp.	Los Algarrobos section		An aulacopleurid
Paciphacops²⁶	P. (Paciphacops) sp.	Sierra de las Minitas	Several specimens	Now known as Echidnops taphomimus¹⁹
	P. sp.	Los Algarrobos section		A phacopid
	P.? sp.	Sierra de las Minitas	One specimen (PULR-127 A-B)	A phacopid
Parabouleia⁶²²	P. eldredgei	Quebrada de Talacasto	9 specimens	A calmoniid
Parabouleia⁶²²	P. sp.	Los Algarrobos section		A calmoniid
Talacastops²²²	T. zarelae	Quebrada de Las Aguaditas Sierra de las Minitas	Multiple specimens	A calmoniid
Tarijactinoides¹⁷	T. jarcasensis	Cerro La Chilca Loma de los Piojos	13 specimens	A calmoniid
Tormesiscus²	T. cf. hildae	Sierra de las Minitas	Partial specimen (PULR-130)	A calmoniid
Unguliproetus²	U.? sp.	Sierra de las Minitas	Two associated fragments	A proetid

References

Cichowolski, Marcela; Rustán, Juan J. (April 2017). "First report of Devonian bactritids (Cephalopoda) from South America: paleobiogeographic and biostratigraphic implications". Journal of Paleontology. 91 (3): 417–433. Bibcode:2017JPal...91..417C. doi:10.1017/jpa.2017.17. hdl:11336/183599. ISSN 0022-3360.
Rustán, Juan J.; Vaccari, N. Emilio; Astini, Ricardo A. (June 2011). "Early Devonian Trilobites from the Sierra De Las Minitas, Northernmost Precordillera (La Rioja Province), Argentina". Ameghiniana. 48 (2): 226–241. doi:10.5710/AMGH.v48i1(335). hdl:11336/53580. ISSN 0002-7014.
García Muro, Victoria J.; Rubinstein, Claudia V.; Rustán, Juan J.; Steemans, Philippe (2018-10-01). "Palynomorphs from the Devonian Talacasto and Punta Negra Formations, Argentinean Precordillera: New biostratigraphic approach". Journal of South American Earth Sciences. 86: 110–126. Bibcode:2018JSAES..86..110G. doi:10.1016/j.jsames.2018.06.009. hdl:11336/87283. ISSN 0895-9811.
Sanchez, Teresa M.; Waisfeld, Beatriz G.; Toro, Blanca A. (1995). "Silurian and Devonian Molluscan Bivalves from Precordillera Region, Western Argentina". Journal of Paleontology. 69 (5): 869–886. Bibcode:1995JPal...69..869S. doi:10.1017/S002233600003554X. ISSN 0022-3360. JSTOR 1306352.
Herrera, Zarela A. (1995). "The Lower Devonian chonetoidean brachiopods from the Argentine Precordillera". Travaux et Documents des Laboratoires de Géologie de Lyon. 136 (1): 101–147.
Carrera, Marcelo G.; Montoya, Ezequiel; Rustán, Juan J.; Halpern, Karen (2013). "Silurian-Devonian coral associations across a sequence stratigraphic boundary in the Argentine Precordillera". Geological Journal. 48 (2–3): 256–269. Bibcode:2013GeolJ..48..256C. doi:10.1002/gj.1332. hdl:11336/24993. ISSN 1099-1034.
Herrera, Zarela Angélica (1995-01-01). "The first notanopliid brachiopod from the South American Devonian sequence". Geobios. 28 (3): 337–342. Bibcode:1995Geobi..28..337H. doi:10.1016/S0016-6995(95)80008-5. ISSN 0016-6995.
Levy, Regina; Nullo, Francisco E. (1972). "Chonetidae Devónicos de Argentina. 1. Chonostrophia". Ameghiniana. 9 (2): 138–144.
Racheboeuf, Patrick R.; Herrera, Zarela (1994-09-01). "On some new Malvinokaffric Silurian and Devonian chonetacean brachiopods, and reclassification of others". Neues Jahrbuch für Geologie und Paläontologie - Monatshefte. 1994 (9): 541–560. doi:10.1127/njgpm/1994/1994/541. ISSN 0028-3630.
Herrera, Zarela Angelica; Salas, María José; Giolitti, Javier A. (January 1998). "Chilidiopsoidea (Brachiopoda) del Devónico Inferior de la Precordillera Argentina". Spanish Journal of Palaeontology (in Spanish). 13 (2): 149–166. doi:10.7203/sjp.24049. ISSN 2660-9568.
García-López, Susana; Fernández Martínez, Esperanza (1995-01-01). "The genus Parastriatopora Sokolov, 1949 (Tabulata) in the Lower Devonian of Argentina: Palaeobiogeographic implications". Geobios. 28 (2): 175–183. Bibcode:1995Geobi..28..175G. doi:10.1016/S0016-6995(95)80223-1. ISSN 0016-6995.
Martínez, Sergio; Río, Claudia del (2015). "Fossil Asterozoa (Echinodermata) of Argentina". Revista de Biología Tropical. 63 (2): 1–6. doi:10.15517/rbt.v63i2.23123 (inactive 12 July 2025).{{cite journal}}: CS1 maint: DOI inactive as of July 2025 (link)
Salas, María José (October 2016). "New Insights on Earliest Devonian (Lochkovian) Ostracods from the Argentine Precordillera". Ameghiniana. 53 (5): 565–585. doi:10.5710/AMGH.04.07.2016.2967. ISSN 0002-7014.
Salas, María J.; Rustán, Juan J.; Sterren, Andrea F. (2013-08-01). "Lower and Middle Devonian Malvinokaffric ostracods from the Precordillera Basin of San Juan, Argentina". Journal of South American Earth Sciences. 45: 56–68. Bibcode:2013JSAES..45...56S. doi:10.1016/j.jsames.2013.02.002. hdl:11336/24991. ISSN 0895-9811.
Carrera, Marcelo G.; Rustán, Juan José (November 2015). "The new genus Talacastospongia: insights on the first record of a Devonian sponge from South America". Journal of Paleontology. 89 (6): 912–919. Bibcode:2015JPal...89..912C. doi:10.1017/jpa.2016.9. hdl:11336/43851. ISSN 0022-3360.
Randolfe, Enrique A.; RustáN, Juan José; Bignon, Arnaud (2022-01-12). "New dalmanitid trilobite taxa from the Lower Devonian of Argentina: comments on spinosity in endemic taxa from southern high paleolatitudes". Comptes Rendus Palevol. 21 (2): 17–38. doi:10.5852/cr-palevol2022v21a2. ISSN 1777-571X.
Vaccari, Norberto E.; Waisfeld, Beatriz G.; Edgecombe, Gregory D. (1994-01-01). "Calmoniid Trilobites of the Lower Devonian Scaphiocoelia zone in the Argentine Precordillera". Geobios. 27 (5): 591–608. Bibcode:1994Geobi..27..591V. doi:10.1016/S0016-6995(94)80257-2. ISSN 0016-6995.
Rustán, Juan José; Vaccari, N. Emilio (February 2012). "A revision of the Devonian Malvinokaffric dalmanitid trilobite Dalmanitoides Delo, 1935, on the basis of new data from Argentina". Palaeontologia Electronica. 15 (1): 1–21. doi:10.26879/276.
Rustán, Juan José; Balseiro, Diego (November 2016). "The phacopid trilobite Echidnops taphomimus n. sp. from the Lower Devonian of Argentina: insights into infaunal molting, eye architecture and geographic distribution". Journal of Paleontology. 90 (6): 1100–1111. Bibcode:2016JPal...90.1100R. doi:10.1017/jpa.2016.108. hdl:11336/44268. ISSN 0022-3360.
Rustán, Juan J.; Vaccari, N. Emilio (2010). "The Aulacopleurid Trilobite Maurotarion Alberti, 1969, in the Silurian-Devonian of Argentina: Systematic, Phylogenetic and Paleobiogeographic Significance". Journal of Paleontology. 84 (6): 1082–1098. Bibcode:2010JPal...84.1082R. doi:10.1666/09-166.1. ISSN 0022-3360. JSTOR 40925985. S2CID 129791708.
Rustán, J.; Vaccari, N. (2012-08-01). "The trilobite Maurotarion megacephalum sp. nov. (Aulacopleuridae) in the Lower Devonian of Argentina: phylogenetic and paleobiogeographic remarks". Revista Mexicana de Ciencias Geológicas. 29: 346–354. S2CID 55274075.
Edgecombe, Gregory D.; Vaccari, Norberto E.; Waisfeld, Beatriz G. (July 1994). "Lower Devonian calmoniid trilobites from the Argentine Precordillera: new taxa of the Bouleia Group, and remarks on the tempo of calmoniid radiation". Geological Magazine. 131 (4): 449–464. Bibcode:1994GeoM..131..449E. doi:10.1017/S0016756800012097. ISSN 1469-5081.

[:0-1] Cichowolski, Marcela; Rustán, Juan J. (April 2017). "First report of Devonian bactritids (Cephalopoda) from South America: paleobiogeographic and biostratigraphic implications". Journal of Paleontology. 91 (3): 417–433. Bibcode:2017JPal...91..417C. doi:10.1017/jpa.2017.17. hdl:11336/183599. ISSN 0022-3360.

[:1-2] Rustán, Juan J.; Vaccari, N. Emilio; Astini, Ricardo A. (June 2011). "Early Devonian Trilobites from the Sierra De Las Minitas, Northernmost Precordillera (La Rioja Province), Argentina". Ameghiniana. 48 (2): 226–241. doi:10.5710/AMGH.v48i1(335). hdl:11336/53580. ISSN 0002-7014.

[3] García Muro, Victoria J.; Rubinstein, Claudia V.; Rustán, Juan J.; Steemans, Philippe (2018-10-01). "Palynomorphs from the Devonian Talacasto and Punta Negra Formations, Argentinean Precordillera: New biostratigraphic approach". Journal of South American Earth Sciences. 86: 110–126. Bibcode:2018JSAES..86..110G. doi:10.1016/j.jsames.2018.06.009. hdl:11336/87283. ISSN 0895-9811.

[:6-4] Sanchez, Teresa M.; Waisfeld, Beatriz G.; Toro, Blanca A. (1995). "Silurian and Devonian Molluscan Bivalves from Precordillera Region, Western Argentina". Journal of Paleontology. 69 (5): 869–886. Bibcode:1995JPal...69..869S. doi:10.1017/S002233600003554X. ISSN 0022-3360. JSTOR 1306352.

[:2-5] Herrera, Zarela A. (1995). "The Lower Devonian chonetoidean brachiopods from the Argentine Precordillera". Travaux et Documents des Laboratoires de Géologie de Lyon. 136 (1): 101–147.

[:7-6] Carrera, Marcelo G.; Montoya, Ezequiel; Rustán, Juan J.; Halpern, Karen (2013). "Silurian-Devonian coral associations across a sequence stratigraphic boundary in the Argentine Precordillera". Geological Journal. 48 (2–3): 256–269. Bibcode:2013GeolJ..48..256C. doi:10.1002/gj.1332. hdl:11336/24993. ISSN 1099-1034.

[7] Herrera, Zarela Angélica (1995-01-01). "The first notanopliid brachiopod from the South American Devonian sequence". Geobios. 28 (3): 337–342. Bibcode:1995Geobi..28..337H. doi:10.1016/S0016-6995(95)80008-5. ISSN 0016-6995.

[:3-8] Levy, Regina; Nullo, Francisco E. (1972). "Chonetidae Devónicos de Argentina. 1. Chonostrophia". Ameghiniana. 9 (2): 138–144.

[:4-9] Racheboeuf, Patrick R.; Herrera, Zarela (1994-09-01). "On some new Malvinokaffric Silurian and Devonian chonetacean brachiopods, and reclassification of others". Neues Jahrbuch für Geologie und Paläontologie - Monatshefte. 1994 (9): 541–560. doi:10.1127/njgpm/1994/1994/541. ISSN 0028-3630.

[:5-10] Herrera, Zarela Angelica; Salas, María José; Giolitti, Javier A. (January 1998). "Chilidiopsoidea (Brachiopoda) del Devónico Inferior de la Precordillera Argentina". Spanish Journal of Palaeontology (in Spanish). 13 (2): 149–166. doi:10.7203/sjp.24049. ISSN 2660-9568.

[11] García-López, Susana; Fernández Martínez, Esperanza (1995-01-01). "The genus Parastriatopora Sokolov, 1949 (Tabulata) in the Lower Devonian of Argentina: Palaeobiogeographic implications". Geobios. 28 (2): 175–183. Bibcode:1995Geobi..28..175G. doi:10.1016/S0016-6995(95)80223-1. ISSN 0016-6995.

[:8-12] Martínez, Sergio; Río, Claudia del (2015). "Fossil Asterozoa (Echinodermata) of Argentina". Revista de Biología Tropical. 63 (2): 1–6. doi:10.15517/rbt.v63i2.23123 (inactive 12 July 2025).{{cite journal}}: CS1 maint: DOI inactive as of July 2025 (link)

[:12-13] Salas, María José (October 2016). "New Insights on Earliest Devonian (Lochkovian) Ostracods from the Argentine Precordillera". Ameghiniana. 53 (5): 565–585. doi:10.5710/AMGH.04.07.2016.2967. ISSN 0002-7014.

[14] Salas, María J.; Rustán, Juan J.; Sterren, Andrea F. (2013-08-01). "Lower and Middle Devonian Malvinokaffric ostracods from the Precordillera Basin of San Juan, Argentina". Journal of South American Earth Sciences. 45: 56–68. Bibcode:2013JSAES..45...56S. doi:10.1016/j.jsames.2013.02.002. hdl:11336/24991. ISSN 0895-9811.

[15] Carrera, Marcelo G.; Rustán, Juan José (November 2015). "The new genus Talacastospongia: insights on the first record of a Devonian sponge from South America". Journal of Paleontology. 89 (6): 912–919. Bibcode:2015JPal...89..912C. doi:10.1017/jpa.2016.9. hdl:11336/43851. ISSN 0022-3360.

[:13-16] Randolfe, Enrique A.; RustáN, Juan José; Bignon, Arnaud (2022-01-12). "New dalmanitid trilobite taxa from the Lower Devonian of Argentina: comments on spinosity in endemic taxa from southern high paleolatitudes". Comptes Rendus Palevol. 21 (2): 17–38. doi:10.5852/cr-palevol2022v21a2. ISSN 1777-571X.

[:9-17] Vaccari, Norberto E.; Waisfeld, Beatriz G.; Edgecombe, Gregory D. (1994-01-01). "Calmoniid Trilobites of the Lower Devonian Scaphiocoelia zone in the Argentine Precordillera". Geobios. 27 (5): 591–608. Bibcode:1994Geobi..27..591V. doi:10.1016/S0016-6995(94)80257-2. ISSN 0016-6995.

[18] Rustán, Juan José; Vaccari, N. Emilio (February 2012). "A revision of the Devonian Malvinokaffric dalmanitid trilobite Dalmanitoides Delo, 1935, on the basis of new data from Argentina". Palaeontologia Electronica. 15 (1): 1–21. doi:10.26879/276.

[:11-19] Rustán, Juan José; Balseiro, Diego (November 2016). "The phacopid trilobite Echidnops taphomimus n. sp. from the Lower Devonian of Argentina: insights into infaunal molting, eye architecture and geographic distribution". Journal of Paleontology. 90 (6): 1100–1111. Bibcode:2016JPal...90.1100R. doi:10.1017/jpa.2016.108. hdl:11336/44268. ISSN 0022-3360.

[20] Rustán, Juan J.; Vaccari, N. Emilio (2010). "The Aulacopleurid Trilobite Maurotarion Alberti, 1969, in the Silurian-Devonian of Argentina: Systematic, Phylogenetic and Paleobiogeographic Significance". Journal of Paleontology. 84 (6): 1082–1098. Bibcode:2010JPal...84.1082R. doi:10.1666/09-166.1. ISSN 0022-3360. JSTOR 40925985. S2CID 129791708.

[21] Rustán, J.; Vaccari, N. (2012-08-01). "The trilobite Maurotarion megacephalum sp. nov. (Aulacopleuridae) in the Lower Devonian of Argentina: phylogenetic and paleobiogeographic remarks". Revista Mexicana de Ciencias Geológicas. 29: 346–354. S2CID 55274075.

[:10-22] Edgecombe, Gregory D.; Vaccari, Norberto E.; Waisfeld, Beatriz G. (July 1994). "Lower Devonian calmoniid trilobites from the Argentine Precordillera: new taxa of the Bouleia Group, and remarks on the tempo of calmoniid radiation". Geological Magazine. 131 (4): 449–464. Bibcode:1994GeoM..131..449E. doi:10.1017/S0016756800012097. ISSN 1469-5081.

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