| Xenoscelinae | |
|---|---|
| |
| Macrophagus robustus | |
Scientific classification 
| |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Clade: | Pancrustacea |
| Class: | Insecta |
| Order: | Coleoptera |
| Suborder: | Polyphaga |
| Infraorder: | Cucujiformia |
| Family: | Erotylidae |
| Subfamily: | Xenoscelinae Ganglbauer, 1899 |
| Synonyms | |
| |
Xenoscelinae is a subfamily of the pleasing fungus beetles (family Erotylidae). They have been recorded from almost all over the world, but most genera are small or monotypic; also, circumscription of this subfamily heavily varies among authors.1
Some define the subfamily more widely, downranking the Cryptophilinae (including Empocryptinae and Toraminae), Loberinae and Pharaxonothinae (including Setariolinae) to tribes of subfamily Xenoscelinae, in which case the core xenoscelines also become a tribe Xenoscelini. Earlier, all these taxa were usually considered to be more closely related to the lizard beetles than to the "typical" pleasing fungus beetles of subfamily Erotylinae, and united with the lizard beetles in a distinct family Languriidae. But the "Languriidae" turned out to be paraphyletic with Erotylidae and eventually were merged into it, leading to a re-sorting of subtaxa that is still not defintely resolved as of 2025.12
Genera
These genera belong to the subfamily Xenoscelinae in its narrow definition (= tribe Xenoscelini, if the Xenoscelinae are defined sensu lato):
- Arrowcryptus Leschen & Węgrzynowicz, 20081
- Leucohimatiops Heller, 19231
- Loberonotha Sen Gupta and Crowson, 196921
- Macrophagus Motschulsky, 18451
- Othniocryptus Sharp, 19001
- Protoloberus Leschen, 200321
- Xenocryptus Arrow, 19291
- Xenoscelis Wollaston, 186421
- Zavaljus Reitter, 1880
The former genus Rhopalocryptus, believed to be a xenosceline pleasing fungus beetle, is actually a narrow-waisted bark beetle (family Salpingidae) of genus Trogocryptoides.
Two fossil genera have been assigned to the Xenoscelinae in its narrow circumscription:3
- †Khasurtyphilus Lyubarsky & Perkovsky, 2025
- †Turgaphilus Lyubarsky & Perkovsky, 2025 (Barremian – Aptian)
References
References
Media related to Xenoscelini at Wikimedia Commons
- Leschen, Richard A.B. & Węgrzynowicz, Piotr (1998): Generic catalogue and taxonomic status of Languriidae (Cucujoidea). Ann.Zool. 48(3/4): 221-243. PDF fulltext
- "Xenoscelinae Ganglbauer, 1899". Global Biodiversity Information Facility. Retrieved 2026-06-23.
- "Xenoscelini Ganglbauer, 1899". BioLib. Retrieved 2026-06-23.
Further reading
Further reading
- Arnett, R.H. Jr.; Thomas, M. C.; Skelley, P. E.; Frank, J. H., eds. (2002). American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press. ISBN 978-0849309540.
- Blatchley, W.S. (1910). An illustrated descriptive catalogue of the Coleoptera, beetles (exclusive of the Rhynchophora) known to occur in Indiana. Nature Pub.
- Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88). Pensoft Publishers: 1–972. doi:10.3897/zookeys.88.807. ISSN 1313-2989. PMC 3088472. PMID 21594053.
- Crotch, G.R. (1873). Check list of the Coleoptera of America, north of Mexico. Naturalists' Agency. doi:10.5962/bhl.title.38811. ISBN 0665070772.
{{cite book}}: ISBN / Date incompatibility (help) - Evans, Arthur V.; Hogue, James N. (2006). Field Guide to Beetles of California. University of California Press. ISBN 978-0520246577.
- Klimaszewski, Jan; Watt, J.Charles (1997). "Coleoptera: family-group review and keys to identification". Fauna of New Zealand. 37.
- Lawrence, J.F.; Newton Jr., A.F. (1995). Pakaluk, James; Slipinski, Stanislaw Adam (eds.). Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson. Vol. 2. Muzeum i Instytut Zoologii PAN. pp. 779–1006. ISBN 83-85192-34-4.
- Lawrence, J.F.; Vaurie, P. (1983). "Family: Languriidae". A Catalog of the Coleoptera of America North of Mexico. United States Department of Agriculture.
- LeConte, J.L. (1861). Classification of the Coleoptera of North America. Smithsonian Miscellaneous Collections. Vol. 3. Smithsonian Institution. doi:10.5962/bhl.title.38459. ISBN 0665100558.
{{cite book}}: ISBN / Date incompatibility (help) - Leng, Charles W. (1920). Catalogue of the Coleoptera of America, North of Mexico. John D. Sherman, Jr. doi:10.5962/bhl.title.8777.
- Leschen, Richard A.B. (2003). "Erotylidae (Insecta: Coleoptera: Cucujoidea) phylogey and review" (PDF). Fauna of New Zealand. 47. Manaaki Whenua Press. ISBN 0-478-09350-0. ISSN 0111-5383. Archived from the original (PDF) on 2019-01-26. Retrieved 2020-05-06.
- Lord, Nathan Patrick; Hartley, Christopher S.; Lawrence, John F.; McHugh, Joseph V.; et al. (2010). "Phylogenetic analysis of the minute brown scavenger beetles (Coleoptera: Latridiidae), and recognition of a new beetle family, Akalyptoischiidae fam. n. (Coleoptera: Cucujoidea)". Systematic Entomology. 35 (4): 753–763. doi:10.1111/j.1365-3113.2010.00532.x. ISSN 0307-6970.
- Poole, Robert W.; Gentili, Patricia, eds. (1996). "A Check List of the Insects of North America: Coleoptera". Nomina Insecta Nearctica: A Check List of the Insects of North America. 1: Coleoptera, Strepsiptera. Entomological Information Services: 41–820. ISBN 9781889002019. Archived from the original on 2020-11-12. Retrieved 2018-04-01.
- Skelley, Paul E.; McHugh, Joseph V. (2002). Arnett Jr., R. Jr.; Thomas, M.; Skelley, P.; Frank, J. (eds.). Family 87. Erotylidae Leach 1815. American Beetles vol. 2, Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press. pp. 348–355. ISBN 0-8493-0954-9.
- White, Richard E. (1998) [1983]. A Field Guide to the Beetles of North America (Peterson Field Guides). Houghton Mifflin Harcourt. ISBN 0395910897.