Article · Wikipedia archive · Last revised Jun 18, 2026

Dinomorpha

Dinomorpha is a proposed clade of extinct therapsids uniting Biarmosuchia and Dinocephalia to the exclusion of other major therapsid groups. The usage of the grouping was introduced in a paper focusing on early therapsid evolution in eastern Europe. Most phylogentic analysis had supported the alternative, more traditional placement of biarmosuchians and dinocephalians in a pectinate arrangement relative to the clade Neotherapsida, a 2026 paper from Duhamel et al. focusing on the phylogenetic relationships of therapsids using exclusively on cranial characters, recovered a clade uniting Biarmosuchia and Dinocephalia. The authors recommended using Ivachnenko (2008) "Dinomorpha" as the available clade name for Biarmosuchia and Dinocephalia, along with using modern definitions for both members in a stem-based manner.

Last revised
Jun 18, 2026
Read time
≈ 1 min
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239 w
Citations
4
Source
Dinomorpha
Temporal range: Middle Permian - Late Permian,
Mounted skeleton of Biarmosuchus tener
Mounted skeleton of Estemmenosuchus uralensis
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Clade: Synapsida
Clade: Therapsida
Clade: Dinomorpha
Ivachnenko, 2008 emmend. Duhamel et al., 2026
Subgroups

Dinomorpha is a proposed clade of extinct therapsids uniting Biarmosuchia and Dinocephalia to the exclusion of other major therapsid groups. The usage of the grouping was introduced in a paper focusing on early therapsid evolution in eastern Europe.1 Most phylogentic analysis had supported the alternative, more traditional placement of biarmosuchians and dinocephalians in a pectinate arrangement relative to the clade Neotherapsida (Anomodontia and Theriodontia),2 a 2026 paper from Duhamel et al. focusing on the phylogenetic relationships of therapsids using exclusively on cranial characters, recovered a clade uniting Biarmosuchia and Dinocephalia.3 The authors recommended using Ivachnenko (2008) "Dinomorpha" as the available clade name for Biarmosuchia and Dinocephalia, along with using modern definitions for both members in a stem-based manner.3

References

References

  1. Ivakhnenko, M.F. Cranial morphology and evolution of Permian Dinomorpha (Eotherapsida) of eastern Europe. Paleontol. J. 42, 859–995 (2008).
  2. Angielczyk, Kenneth D.; Kammerer, Christian F. (2018-10-22). "Non-Mammalian synapsids: the deep roots of the mammalian family tree". In Zachos, Frank; Asher, Robert (eds.). Mammalian Evolution, Diversity and Systematics. De Gruyter. pp. 117–198. doi:10.1515/9783110341553-005. ISBN 978-3-11-034155-3. S2CID 92370138.
  3. Duhamel, A.; Wynd, B.; Wright, A. M.; Moopen, A.; Benoit, J.; Rubidge, B. (2026). "Rethinking therapsid phylogeny through Bayesian and cladistic approaches". Scientific Reports. doi:10.1038/s41598-026-38195-2. PMID 41813723.