Corydalidae

Corydalidae Temporal range:
a Corydalus species
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Clade:	Pancrustacea
Class:	Insecta
Order:	Megaloptera
Family:	Corydalidae Leach 1815
Subfamilies
Chauliodinae Corydalinae

The family Corydalidae contains the megalopterous insects known as dobsonflies and fishflies. Making up about three dozen genera,¹ they occur primarily throughout North America, both temperate and tropical, South America, Australia, New Zealand, Africa (particularly South Africa) and Asia.

They are sizeable Megaloptera, with a body usually larger than 25 mm (1 inch). They often have long filamentous antennae, though in male fishflies they are characteristically feathered. Ocelli are present; the fourth tarsal segment is cylinder-shaped. The four large wings are translucent, smoky grey, or mixed, and the anterior pair is slightly longer than the posterior one. Their aquatic larvae are used as fish bait and are called hellgrammites. The larvae are aquatic, active, armed with strong sharp mandibles, and breathe by means of abdominal branchial filaments. When full sized — which can take several years — they leave the water and spend a quiescent pupal stage on the land, in chambers dug under stones or logs, before metamorphosis into the sexually mature insect.

• 
  Parachauliodes japonicus
• 
  Larva

The eastern dobsonfly, Corydalus cornutus, is the most well-known North American species among the dobsonflies. These genera have distinctive elongated mandibles in males and form the subfamily Corydalinae. The genera in which the males have normal mandibles, called fishflies, form the subfamily Chauliodinae. The summer fishfly, Chauliodes pectinicornis, is perhaps the best-known of these in North America; its immense mating swarms in the Upper Mississippi River region fill the air on a few summer nights each year much like mayflies in certain regions of Europe, leaving millions of carcasses to be cleaned up the next day.

• 
  Chauliodes pectinicornis
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  Acanthacorydalis asiatica

The family is divided into two subfamilies, the Corydalinae and the Chauliodinae. Classification and identification is based on external morphology and at the species level using male genitalia.² These 31 genera belong to the family Corydalidae as of 2022:³