| Copiula | |
|---|---|
Scientific classification 
| |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Amphibia |
| Order: | Anura |
| Family: | Microhylidae |
| Subfamily: | Asterophryinae |
| Genus: | Copiula Méhelÿ, 1901 |
| Type species | |
| Phrynixalus oxyrhinus Boulenger, 1898
| |
| Species | |
|
14 species (see text) | |
Copiula is a genus of microhylid frogs endemic to New Guinea. The common name Mehely frogs has been coined for them.1 They are leaf-litter inhabitants.2
Taxonomy
Copiula is probably not monophyletic. Some former Austrochaperina species have already been transferred to this genus, and further ones might follow when more data became available.13
Species
There are at present 14 species in this genus:1
- Copiula alpestris (Zweifel, 2000)
- Copiula annanoreenae Günther, Richards, and Dahl, 2014
- Copiula derongo (Zweifel, 2000)
- Copiula exspectata Günther, 2002
- Copiula fistulans Menzies and Tyler, 1977 — Lae Mehely Frog
- Copiula guttata (Zweifel, 2000)
- Copiula lennarti Günther, Richards, and Dahl, 2014
- Copiula major Günther, 2002
- Copiula minor Menzies and Tyler, 1977 — Milne Bay Mehely Frog
- Copiula obsti Günther, 2002
- Copiula oxyrhina (Boulenger, 1898) — Misima Island Mehely Frog
- Copiula pipiens Burton and Stocks, 1986 — Wirui Mehely Frog
- Copiula rivularis (Zweifel, 2000)
- Copiula tyleri Burton, 1990 — Sepik Mehely Frog
The AmphibiaWeb4 reports fewer species, with species that Peloso and colleagues moved in 2016 from Austrochaperina and Oxydactyla missing.3
References
References
- Frost, Darrel R. (2016). "Copiula Méhely, 1901". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 20 July 2016.
- Zweifel, R. G. (2000). "Partition of the Australopapuan microhylid frog genus Sphenophryne with descriptions of new species". Bulletin of the American Museum of Natural History. 253: 1–130. doi:10.1206/0003-0090(2000)253<0001:POTAMF>2.0.CO;2. hdl:2246/1600. S2CID 85621508.
- Peloso, Pedro L.V.; Frost, Darrel R.; Richards, Stephen J.; Rodrigues, Miguel T.; Donnellan, Stephen; Matsui, Masafumi; Raxworthy, Cristopher J.; Biju, S.D.; Lemmon, Emily Moriarty; Lemmon, Alan R.; Wheeler, Ward C. (2016). "The impact of anchored phylogenomics and taxon sampling on phylogenetic inference in narrow-mouthed frogs (Anura, Microhylidae)". Cladistics. 32 (2): 113–140. doi:10.1111/cla.12118. PMID 34732021. S2CID 84925667.
- "Microhylidae". AmphibiaWeb: Information on amphibian biology and conservation. [web application]. Berkeley, California: AmphibiaWeb. 2016. Retrieved 20 July 2016.