Article · Wikipedia archive · Last revised Jun 19, 2026

Chorrillo Formation

The Chorrillo Formation, also named as Chorillo Formation, is a Maastrichtian geologic formation in southern Patagonia, Argentina. The formation is more than 50 metres (160 ft) thick and underlies the Calafate Formation and rests on top of the La Irene Formation. It constitutes a significant uppermost Cretaceous continental sedimentary sequence within the Magallanes Basin, located in Santa Cruz Province, Argentina, approximately 30 km southwest of El Calafate. Initial investigations in the 1990s laid the groundwork, with detailed paleontological studies emerging since 2019, revealing a rich biotic assemblage. The formation correlates with the lower Dorotea Formation in Chile, sharing dinosaurian ecotypes, while both are clearly different from other northern Patagonian Maastrichtian units, which are dominated by saltasaurine sauropods and abelisaurids, suggesting potential temporal or environmental distinctions.

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Jun 19, 2026
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Chorrillo Formation
Stratigraphic range:
Satellite image of the Chorrillo Formation site (in the middle on the right). In the bottom left is the Perito Moreno Glacier.
TypeGeological formation
UnderliesCalafate Formation
OverliesLa Irene Formation
Thickness>50 m (160 ft)
Lithology
PrimaryClaystone
Location
LocationPatagonia
Coordinates49°54′S 72°30′W / 49.9°S 72.5°W / -49.9; -72.5
Approximate paleocoordinates51°48′S 62°12′W / 51.8°S 62.2°W / -51.8; -62.2
RegionSanta Cruz Province
CountryArgentina
ExtentSoutheast of Viedma Lake
North of Argentino Lake
Austral Basin
Type section
Named byArbe & Hechem
Year defined1984
Chorrillo Formation (Argentina)

The Chorrillo Formation, also named as Chorillo Formation,1 is a Maastrichtian (Late Cretaceous Epoch, 72.1–66 million years ago) geologic formation in southern Patagonia, Argentina. The formation is more than 50 metres (160 ft) thick and underlies the Calafate Formation and rests on top of the La Irene Formation.23 It constitutes a significant uppermost Cretaceous continental sedimentary sequence within the Magallanes Basin, located in Santa Cruz Province, Argentina, approximately 30 km southwest of El Calafate. Initial investigations in the 1990s laid the groundwork, with detailed paleontological studies emerging since 2019, revealing a rich biotic assemblage.4 The formation correlates with the lower Dorotea Formation in Chile, sharing dinosaurian ecotypes, while both are clearly different from other northern Patagonian Maastrichtian units (e.g., Allen Formation, Los Alamitos Formation), which are dominated by saltasaurine sauropods and abelisaurids, suggesting potential temporal or environmental distinctions.34

Geology

The Chorrillo Formation is embedded within the foreland Magallanes Basin, shaped by Late Cretaceous Andean orogeny driven by subduction along the Pacific margin. Tectonic subsidence, coupled with sediment supply from the rising Patagonian Andes, facilitated deposition.4 Lithologically, the unit comprises fine- to coarse-grained clastics: mudstones and siltstones dominate floodplain facies, often organic-rich and pedogenically modified; sandstones, arkosic to lithic, form channel fills with cross-bedding; and conglomerates appear as basal lags.5 Clay mineralogy is dominated by smectite, with minor illite and kaolinite, reflecting volcanic ash alteration. Sedimentologically, it records a meandering fluvial system on a distal floodplain, featuring fining-upward channel sequences, crevasse splays, and overbank deposits.3 High-frequency paleosol stacking indicates local hydrologic and topographic variability, while low-frequency patterns suggest avulsion-driven progradation. Regional ties extend to the Dorotea Formation in Chile, reflecting basin-wide tectonic influences.5

Paleoenvironment

Modern equivalent, Valdivian Rainforest source ↗

Paleoenvironmental reconstructions indicate a temperate-warm, seasonally humid climate with mean annual temperatures of 10-18 °C and precipitation of 900–1500 mm, inferred from paleosols, clay mineralogy, and biotic indicators.6 Depositional settings encompass low-energy fluvial floodplains with paludal and lacustrine elements, marked by poor drainage, hydromorphic paleosols (e.g., vertisols, histosols), and redoximorphic features such as mottling and iron nodules, suggesting fluctuating water tables and episodic anoxia.5 Aquatic communities feature Nymphaeaceae, Salviniaceae, Salviniaceae, and Zygnemataceae, indicative of freshwater ponds and swamps, while terrestrial elements include conifers, ferns, and angiosperms in riparian zones.7 Floral phytozones, derived from palynological data, comprise five assemblages across the section, exhibiting quantitative variations in taxa proportions without major turnover: lower zones are fern-dominated (e.g., Cyatheaceae), transitioning to angiosperm-rich upper zones (e.g., Proteaceae, Nothofagidites), attributed to subtle humidity fluctuations or habitat gradients relative to fluvial channels.8 These patterns align with Maastrichtian assemblages in southern Gondwanan basins (e.g., La Anita, Lefipán Formation), reflecting angiosperm diversification and Weddellian Province expansion.38

Fossils

Dinosaur remains are among the fossils that have been recovered from the formation.

Fossils recovered from the formation are:

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Arthropoda

Decapoda

Genus Species Locality Material Notes Images
Astacidea9 Indeterminate "Locality 4" MPM 21516, three gastroliths Previously identified as crushing fish teeth
Extant example source ↗

Insects

Genus Species Locality Material Notes Images
Coelolepida10 Indeterminate La Anita farm MPM-Pal 21835-3:W34/1; MPM-Pal 21835-23:W55/0; MPM-Pal 21835-22:R52/2; MPM-Pal 21835-20:Z47/3; MPM-Pal 21835-22: G55/0; MPM-Pal 2183-21: E42/3 A Glossatan Lepidopteran
Diamesinae10 "Morphotype 3" La Anita farm MPM-Pal 21835-9:E34/4; MPM-Pal 21835-19: V34/4 A Chironomidae Nematoceran. Similar to the living genus Paraheptagyia
Extant example source ↗
Ephemeroptera10 Indeterminate La Anita farm MPM-Pal 21835-9:W37/1 A Mayfly
Lepidoptera10 Indeterminate La Anita farm MPM-Pal 21835-10:X30/2; MPM-Pal 21835-7:S27/2
Orthocladiinae10 "Morphotype 1" La Anita farm MPM-Pal 21835-24: M31/0; MPM-Pal 21835-3: P46/4 A Chironomidae Nematoceran
"Morphotype 2" La Anita farm MPM-Pal 21835-25: V26/3
"Morphotype 5" La Anita farm MPM-Pal 21835-9: Q58/3 It resembles modern Cricotopus
Cricotopus source ↗
Tanypodinae10 "Morphotype 4" La Anita farm MPM-Pal 21835-2:Y40/3 A Chironomidae Nematoceran
Extant example source ↗

Molluscs

Gastropods

Genus Species Locality Material Notes Images
Allopeas11 A. agnolini "Locality 4" Shells An achatinid gastropod
Extant specimen source ↗
Bocourtia11 B. leonardodavincii "Locality 4" Shells A bulimulid gastropod
Extant specimen source ↗
Coelostemma11 C. patagonica "Locality 4" Shells A holospirid gastropod
Holospira4 H. sp. "Locality 4" Shells An urocoptidae gastropod
Leptinaria11 Cf.L. sp. "Locality 4" Shells An achatinid gastropod
Extant specimen source ↗
Lilloiconcha11 L. novasi "Locality 4" Shells A charopid gastropod
Naesiotus11 Cf.N. sp. "Locality 4" Shells A bulimulid gastropod
Extant specimen source ↗
Physa4 P. sp. "Locality 4" Shells A physidae gastropod
Extant specimen source ↗
Physella11 Cf.P. sp. "Locality 4" Shells A physidae gastropod
Extant specimen source ↗
Pleuroceridae4 Indeterminate "Locality 4" Shells Incertade Sedis
Pomacea4 P. sp. "Locality 4" Shells An ampullariidae gastropod
Extant specimen source ↗
Potamolithus11 P. fabiani "Locality 4" Shells A tateid gastropod
Extant specimen source ↗
Stenophysa4 S. sp. "Locality 4" Shells A physidae gastropod

Bivalves

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Genus Species Locality Material Notes Images
Chlamys4 "C." salamanca "Locality 5" Shells A Pectinidae bivalve
Extant specimen source ↗
Cubitostrea4 C. ameghinoi "Locality 5" Shells An Ostreidae bivalve
Extant specimen source ↗
Gryphaeostrea4 G. callophyla "Locality 5" Shells A Gryphaeidae bivalve

Fishes

Genus Species Locality Material Notes Images
Lamniformes4 Indeterminate "Locality 5" Isolated upper lateral tooth Similar to Cretalamna.3
Vidalamiinae4 Indeterminate "Locality 2" Isolated tooth An amiid fish

Amphibians

Genus Species Locality Material Notes Images
Anura4 Indeterminate "Locality 4" MPM 21518, isolated tibiofibula
Calyptocephalella34 C. gayi "Locality 2" MPM-PV-22844, proximal right radioulna; MPM-PV-22845, left proximal end of tibiofibula; MPM-PV-22846, proximal half of urostyle. Calyptocephalellidae frog
Extant specimen of C. gayi source ↗
C. sp. Magallanodon Site maxillae MPM-PV-22841 and MPM-PV-22842; ilium MPM-PV-22843 A species very small in size, distinct from C. gayi.
Calyptocephalellidae4 Indeterminate "Locality 4" MPM 21519, distal end of right humerus with eroded distal margin
Pipoidea34 Indeterminate "Locality 2" Distal end of right humerus Resembles, yet is clearly different, from Xenopus.3

Plesiosaurs

Genus Species Locality Material Notes Images
Elasmosauridae3 Indeterminate "Locality 5" Incomplete cervical centrum and the lateral half of a very fragmentary dorsal vertebra May belong to the Calafate Formation

Lepidosaurs

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Mosasaurs

Genus Species Locality Material Notes Images
Mosasauridae34 Indeterminate "Locality 5" Seven teeth Its presence suggest marine influences

Rhynchocephalian

Genus Species Locality Material Notes Images
Notosphenos12 N. finisterre "Monotreme site" Incomplete right maxilla preserving two completeand one partial teeth. A sphenodontid rhynchocephalian

Snakes

Genus Species Locality Material Notes Images
Alethinophidia412 Indeterminate "Locality 4" MPM 21522 small vertebra
Anilioidea3 Indeterminate "Locality 2" MPM-PV-22847, possible mid-precloacal vertebra
Coniophiidae12 Indeterminate "Monotreme site" Nearly complete mid-posterior precloacal vertebra; Incomplete posterior precloacal vertebra Shares several features with Coniophis.
Madtsoiidae12 Indeterminate "Monotreme site"; "Green Valley site" Incomplete mid-posterior precloacal vertebralacking most of the neural arch; Poorly preserved precloacal vertebra preserving the verteb-ral centrum
Rionegrophis4 R. madtsoioides "Locality 2" A partial vertebra. A madtsoiid snake

Turtles

Genus Species Locality Material Notes Images
Chelidae4 Indeterminate "Locality 2" MPM 21520, two peripheral and one costal plates Largest plates suggest a maximum carapace length of 40 cm.
Hydromedusinae3 Indeterminate "Locality 2, 4, 5" Left peripheral 3 or 4, bridge peripheral, fragment indet; bridge peripheral, right hypoplastron, buttress, plastral fragment and ?plastral fragment Similar to Hydromedusa.
Modern Hydromedusa source ↗
Meiolaniformes3 Indeterminate "Locality 4" Distal end of a left humerus Southernmost record for the clade in South America

Archosaurs

Crocodylomorphs

Genus Species Locality Material Notes Images
Kostensuchus13 K. atrox Estancia La Anita Skull, jaws, incomplete postcranial skeleton A large-bodied, broad-snouted peirosaurid
source ↗

Ornithischians

Genus Species Locality Material Notes Images
Hadrosauridae414 Indeterminate La Anita Farm Two incomplete caudal centra
Isasicursor4 I. santacrucensis "Locality 5" Multiple specimens including teeth, several vertebrae, humerus, thigh bones, shoulder blades, and limbs An elasmarian ornithopod
Isasicursor skeletal source ↗
Ornithopoda414 Indeterminate La Anita Farm Proximal end of a femur A small sized ornithopod
Parankylosauria14 Indeterminate La Anita Farm Five fragmentary caudal vertebrae, incomplete ribs, two incomplete humeri, and nine osteoderms

Sauropods

Genus Species Locality Material Notes Images
Fusioolithus4 F. sp. "Locality 2" Eggshells Fusioolithidae oogenus
Lithostrotia15 Indeterminate Río Leona MACN 5205 (3 middle–posterior caudal centra)
Nullotitan4 N. glaciaris Estancia Alta Vista; "Locality 1, 2, 4, 5" A partial skeleton consisting of a third cervical vertebra, tail vertebrae, a neck rib, ribs, a left shoulder blade, the ends of a right thigh bone, a right shin, a right calf bone, and a right ankle bone A colossosaurian titanosaur framless
Titanosauria4 Indeterminate "Locality 2, 4" Seven isolated teeth4

Theropods

Genus Species Locality Material Notes
Enantiornithes3 Indeterminate "Locality 2, 4" Distal fragment of a pedal phalanx; Two isolated ungual pedal phalanges
Kank16 K. australis "Monotreme site" Fragmentary remains, including vertebrae, teeth, and pedal phalanges A unenlagiid theropod
source ↗
Kookne4 K. yeutensis "Locality 2" An incomplete right coracoid, lacking the sternal end and the proximal end damaged An ornithuran bird similar to Anseriformes
Maip17 M. macrothorax La Anita Farm Axis, several dorsal and caudal vertebrae, cervical and dorsal ribs, gastralia, a left coracoid, fragmentary scapula, partial right pubis, and partial metatarsal The largest megaraptorid theropod of South America and possibly the largest member of this group
M. macrothorax source ↗
Megaraptoridae4 Indeterminate La Anita Farm Isolated teeth4
Noasauridae4 Indeterminate "Locality 4" Isolated right pedal phalanx IV-2?4 Similar to Velocisaurus and Vespersaurus
Prismatoolithidae4 Indeterminate "Locality 4" Eggshells Theropod eggshells
Unenlagiidae4 Indeterminate "Locality 4" An isolated dorsal vertebra, pedal ungual of digit II and phalanx III-24 Similar to Neuquenraptor
Yatenavis18 Y. ieujensis Estancia La Anita Distal right humerus An enantiornithine bird
Yatenavis humerus source ↗

Mammals

Genus Species Locality Material Notes Images
Magallanodon19 M. baikashkenke20 Río de Las Chinas Valley and La Anita Farm Tentative first upper molariform, anterior end of a left dentary bearing part of its gliriform incisor, and upper mesial incisor from the right side A ferugliotheriid gondwanathere
M. terrwerr21 "Locality 2" MPM-PV-23088, isolated right lower molariform; MPM-PV-23089, isolated right lower molariform A ferugliotheriid gondwanathere
Mammalia4 Indeterminate "Locality 4" Two isolated caudal vertebrae Characters shared with mammals having long and gracile tails.
Meridiolestida21 Indeterminate "Monotreme site" MPM-PV-21707, two fragments of a right dentary Small sized taxon similar to Reigitherium.
Mesungulatoidea21 Indeterminate La Anita Farm MPM-PV-21708, fragment of left dentary with five alveoli corresponding with three teeth. Larger than any known Mesozoic meridiolestidan.21
Orretherium21 O. tzei "Monotreme site" MPM-PV-23514, left upper second molariform. A member of Meridiolestida belonging to the family Mesungulatidae.
Patagorhynchus22 P. pascuali "Monotreme site" Lower right molar attached to a fragment of the dentary A ornithorhynchid monotreme
Patagorhynchus holotype source ↗
Patagomaia2123 P. chainko La Anita Farm Partial left acetabulum and ischium; MPM-PV-23515, one fragment of thoracic vertebra, posterior half of one caudal vertebra and 38 indeterminate bones. A large therian
Patagomaia holotype source ↗

Flora

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Algae

Genus Species Affinity / Family Image
Botryococcus68 B. sp. Botryococcaceae
Cymatiosphaera68 C. sp. Zygnemataceae
Leiosphaeridia68 L. sp. Zygnemataceae
Ovoidites68 O. parvus Zygnemataceae
O. spriggii Zygnemataceae
Schizosporis68 S. reticulatus Zygnemataceae (aff. Spirogyra)
Sphaeroplea8 S. striatocristata Sphaeropleaceae

Liverworts

Genus Species Affinity / Family Image
Aequitriradites68 A. spinulosus Marchantiidae
Stoverisporites68 S. microverrucatus Marchantiidae
Taurocusporites68 T. segmentatus Marchantiidae
Triporoletes68 T. reticulatus Marchantiidae
T. sp. Marchantiidae

Hornworts

Genus Species Affinity / Family Image
Foraminisporis68 F. wonthaggiensis Anthocerotidae
F. sp. Anthocerotidae

Lycopods

Genus Species Affinity / Family Image
Camarozonosporites68 C. ambigens Selaginellales
Ceratosporites68 C. equalis Selaginellales
Densoisporites68 D. velatus Selaginellales
Foveosporites68 F. sp. Selaginellales
Retitriletes68 R. austroclavatidites Lycopodiales
R. sp. cf. R. douglasii Lycopodiales

Ferns

Genus Species Affinity / Family Image
Azolla68 A. cf. andreisii Salviniaceae
A. sp. Salviniaceae
Baculatisporites68 B. comaumensis Osmundaceae
B. kachaikensis Osmundaceae
B. turbioensis Osmundaceae
cf. Cladophlebis7 C. sp. Polypodiopsida
Clavifera68 C. sp. Gleicheniaceae
Cyatheacidites68 C. annulatus Dicksoniaceae (aff. Lophosoria)
Cyathidites68 C. australis Cyatheaceae
C. minor Cyatheaceae
C. rafaeli Cyatheaceae
Dictyophyllidites68 D. sp. cf. D. concavus Dipteridaceae / Matoniaceae
Gleicheniidites68 G. sp. Gleicheniaceae
Leptolepidites68 L. verrucatus Blechnaceae
Ornamentifera68 O. sp. Gleicheniaceae
Osmundacidites68 O. diazii Osmundaceae
Peromonolites68 P. vellosus Blechnaceae

Non-conifer Seed Plants

Genus Species Affinity / Family Image
Cycadopites68 C. follicularis Cycadales, Bennettitales, or Ginkgoales
Ephedripites68 E. ambiguous Ephedrales
Equisetosporites68 E. notensis Ephedrales
E. sp. cf. E. dudarensis Ephedrales
Vitreisporites68 V. signatus Pteridosperms

Conifers

Genus Species Affinity / Family Image
Araucariacites68 A. australis Araucariaceae
Callialasporites68 C. turbatus Araucariaceae
Classopollis68 C. sp. Cheirolepidiaceae
Cyclusphaera68 C. psilata Araucariaceae
Dacrycarpites68 D. australiensis Podocarpaceae
Dilwynites68 D. tuberculatus Araucariaceae
Gamerroites68 G. psilasaccus Podocarpaceae
Lygistepollenites68 L. florinii Podocarpaceae
Microcachryidites68 M. antarcticus Podocarpaceae (aff. Microcachrys)
Microcachrys source ↗
Peripollenites68 P. sp. Cupressaceae
Phyllocladidites68 P. mawsonii Podocarpaceae (aff. Lagarostrobos franklinii)
Lagarostrobos source ↗
Podocarpidites68 P. elegans Podocarpaceae
P. elipticus Podocarpaceae
P. exiguus Podocarpaceae
P. herbstii Podocarpaceae
P. major Podocarpaceae
P. marwikii Podocarpaceae
P. otagoensis Podocarpaceae
P. sp. cf. P. langii Podocarpaceae
P. sp. cf. P. microreticuloidata Podocarpaceae
P. torquatus Podocarpaceae
P. verrucosus Podocarpaceae
Podocarpoxylon4 P. dusenii Podocarpaceae
Trichotomosulcites68 T. subgranulatus Podocarpaceae

Magnoliids

Genus Species Affinity / Family Image
Clavatimonocolpites68 C. sp. Magnoliids

Monocots

Genus Species Affinity / Family Image
Arecipites68 A. minutiscabratus Arecaceae
A. sp. cf. A. subverrucatus Arecaceae
Aff.Eolirion7 "Morphotype 5" Palmae
Longapertites68 L. proxapertitoides Arecaceae
Liliacidites68 L. lanceolatus Liliales (aff. Lilium)
Lilium source ↗
L. variegatus Liliales (aff. Lilium)
L. vermireticulatus Liliales
L. sp. cf. L. kaitangataensis Liliaceae/Amaryllidaceae
Monocotyledonean7 "Morphotype 6" Monocots
Nuphar7 N.? sp. 1 Nymphaeaceae
Extant specimen source ↗
N.? sp. 2 Nymphaeaceae
Pandaniidites68 P. typicus Araceae
Proxapertites68 P. operculatus Araceae
Spinozonocolpites68 S. hialinus Arecaceae

Eudicots

Genus Species Affinity / Family Image
Beauprea68 B. elegansiformis Proteales
Beauprea drawing source ↗
Aff. Brachychiton7 "Morphotype 4" Malvaceae
Extant specimen source ↗
Buttinia68 B. andreevii Buxales
Clavamonocolpites68 C. sp. ?Chloranthaceae
Coccoloba?7 "Morphotype 1" Polygonaceae
Extant specimen source ↗
Constantinisporis68 C. jacquei ?Chloranthaceae
Dicotetradites68 D. sp. Ericales
Diporites68 D. sp. ?Chloranthaceae
Doyleipollenites68 D. sp. ?Chloranthaceae
Ericipites68 E. scabratus Ericales
Eudicots7 "Morphotype 3" Eudicots
Myrtaceidites68 M. sp. cf. M. mesonesus Myrtales
Nothofagidites68 N. saraensis Nothofagaceae (cf. Nothofagus)
Nothofagus source ↗
Nyssapollenites68 N. sp. ?Chloranthaceae
Peninsulapollis68 P. gillii Proteaceae (cf. Beauprea?)
P. truswelliae Proteaceae (cf. Beauprea?)
Polycolpites68 P. sp. Pedaliaceae (?cf. Josephinia)
Proteacidites68 P. parvus Proteaceae (cf. Bellendena montana)
Bellendena source ↗
P. subscabratus Proteaceae
P. sp. cf. P. tenuiexinus Proteaceae
P. sp. Proteaceae
Psilatricolporites68 P. sp. cf. P. complanatius Tetrameristaceae
P. sp. Tetrameristaceae
Retidiporites68 R. camachoi Proteaceae
Rhoipites68 R. baculatus Ericales
R. minusculus Ericales
R. sp. cf. R. alveolatus Ericales
R. sp. 1 Ericales
R. sp. 2 Ericales
Rousea68 R. patagonica Ericales
R. sp. cf. R. minuscula Ericales
R. sp. Ericales
Syncolporites68 S. sp. Ericales
Tetracolpites68 T. sp. ?Chloranthaceae
Triatriopollenites68 T. lateflexus Proteaceae
Tricesticillus68 T. americanus ?Chloranthaceae
Tricolpites68 T. reticulatus Gunneraceae (Gunnera macrophylla)
Gunnera source ↗
T. sp. cf. T. brevicolpus Gunneraceae
T. sp. cf. T. inargutus Gunneraceae
T. sp. cf. T. lumbrerensis Gunneraceae
T. sp. 1 Ranunculaceae (cf. Clematis)
Clematis source ↗
Tricolporites68 T. sp. 2 Ericales
T. sp. 3 Ericales
Tricolporoidites68 T. sp. cf. T. eximius Ericales
Triorites68 T. sp. cf. T. minusculus ?Chloranthaceae
Tubulifloridites68 T. lilliei Ericales
Volkheimerites68 V. labyrinthus ?Chloranthaceae
See also

See also

References

References

  1. Río Leona at Fossilworks.org
  2. Marenssi et al., 2003, p.406
  3. Moyano-Paz, D.; Rozadilla, S.; Agnolín, F.; Vera, E.; Coronel, M. D.; Varela, A. N.; Gómez-Dacal, A. R.; Aranciaga-Rolando, A. M.; D'Angelo, J.; Pérez-Loinaze, V.; Richiano, S.; Chimento, N.; Motta, M. J.; Sterli, J.; Manabe, M. (2022-02-01). "The uppermost Cretaceous continental deposits at the southern end of Patagonia, the Chorrillo Formation case study (Austral-Magallanes Basin): Sedimentology, fossil content and regional implications". Cretaceous Research. 130 105059. doi:10.1016/j.cretres.2021.105059. ISSN 0195-6671.
  4. Fernando Novas; Federico Agnolin; Sebastián Rozadilla; Alexis Aranciaga-Rolando; Federico Brissón-Eli; Matias Motta; Mauricio Cerroni; Martín Ezcurra; Agustin Martinelli; Julia D'Angelo; Gerardo Álvarez-Herrera; Adriel Gentil; Sergio Bogan; Nicolas Chimento; Jordi García-Marsà; Gastón Lo Coco; Sergio Miquel; Fatima Brito; Ezequiel Vera; Valeria Loinaze; Mariela Fernandez; Leonardo Salgado (2019). "Paleontological discoveries in the Chorrillo Formation (upper Campanian-lower Maastrichtian, Upper Cretaceous), Santa Cruz Province, Patagonia, Argentina". Revista del Museo Argentino de Ciencias Naturales. Nueva Series. 21 (2): 217–293. doi:10.22179/revmacn.21.655. hdl:11336/122229.
  5. Raigemborn, M. Sol; Lizzoli, Sabrina; Moyano-Paz, Damián; Varela, Augusto N.; Poiré, Daniel G.; Perez Loinaze, Valeria; Vera, Ezequiel; Manabe, Makoto; Novas, Fernando (2023-10-01). "Paleosols of the Maastrichtian dinosaur-bearing Chorrillo Formation (southern Patagonia, Argentina): Paleoenvironmental and paleoclimate implications". Cretaceous Research. 150 105587. doi:10.1016/j.cretres.2023.105587. ISSN 0195-6671.
  6. Perez Loinaze, Valeria S.; Vera, Ezequiel I.; Raigemborn, M. Sol; Moyano-Paz, Damián; Santamarina, Patricio; Agnolín, Federico; Manabe, Makoto; Tsuihiji, Takanobu; Novas, Fernando E. (2025-01-25). "Palynofloras from the Maastrichtian Chorrillo Formation (Patagonia, Argentina): Paleoenvironmental and paleoclimatic implications". Palaeogeography, Palaeoclimatology, Palaeoecology. 662 (in press) 112764. doi:10.1016/j.palaeo.2025.112764. ISSN 0031-0182.
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