Article · Wikipedia archive · Last revised Jun 30, 2026

Tolypelepidida

Tolypelepidida is an extinct order of heterostracan vertebrates. These armored jawless fish superficially resemble their relatives, the cyathaspids, though, researchers place the tolypelepids as a sister group to the cyathaspids and the pteraspidids. A recent study by Lundgren and Blom in 2013 implies that the order is paraphyletic, with the type genus, Tolypelepis, being the sister taxon of Cyathaspidiformes. The typical tolypelepid had a carapace formed from dorsal and ventral plates, and a scaly tail.

Last revised
Jun 30, 2026
Read time
≈ 1 min
Length
167 w
Citations
2
Source
Tolypelepidida
Temporal range: Silurian - Early Devonian
The type species, Tolypelepis undulata
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Infraphylum: Agnatha
Class: Pteraspidomorpha
Subclass: Heterostraci
Order: Tolypelepidida
Denison 1964
Family: Tolypelepididae
Kiær 1932
Genera
  • Asketaspis
  • Athenaegis
  • Tolypelepis
Synonyms
  • Tolypelepidiformes

Tolypelepidida is an extinct order of heterostracan vertebrates. These armored jawless fish superficially resemble their relatives, the cyathaspids, though, researchers place the tolypelepids as a sister group to the cyathaspids and the pteraspidids (and both groups' daughter taxa).1 A recent study by Lundgren and Blom in 2013 implies that the order is paraphyletic, with the type genus, Tolypelepis, being the sister taxon of Cyathaspidiformes.2 The typical tolypelepid had a carapace formed from dorsal and ventral plates, and a scaly tail.

Athenaegis chattertoni cast, middle Silurian, Road River Formation, Mackenzie Mountains, Northwest Territories (Canada). source ↗
Athenaegis chattertoni reconstruction. At the Museo delle Scienze (MUSE). source ↗
References

References

  1. Janvier, Philippe (1997) Heterostraci The Tree of Life Web Project.
  2. Lundgren, Mette, and Henning Blom. "Phylogenetic relationships of the cyathaspidids (Heterostraci)." GFF 135.1 (2013): 74-84.