Article · Wikipedia archive · Last revised Jul 27, 2026

Cyrtomium

Cyrtomium is a genus of about 35 species of ferns in the family Dryopteridaceae, subfamily Dryopteridoideae, according to the Pteridophyte Phylogeny Group classification of 2016. Species are native to Asia, Africa, and the Pacific Ocean islands (Hawaii). It is very closely related to the genus Polystichum, with 2016 research suggesting it should be included in a clade sister to Polystichum s.s.

Last revised
Jul 27, 2026
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Cyrtomium
Cyrtomium falcatum
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Embryophytes
Clade: Tracheophytes
Division: Polypodiophyta
Class: Polypodiopsida
Order: Polypodiales
Suborder: Polypodiineae
Family: Dryopteridaceae
Subfamily: Dryopteridoideae
Genus: Cyrtomium
C.Presl1
Species

See text.

Cyrtomium is a genus of about 35 species of ferns in the family Dryopteridaceae, subfamily Dryopteridoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I).2 Species are native to Asia, Africa (including Madagascar), and the Pacific Ocean islands (Hawaii). It is very closely related to the genus Polystichum, with 2016 research suggesting it should be included in a clade sister to Polystichum s.s.3

Species

As of February 2020, the Checklist of Ferns and Lycophytes of the World accepted the following species:1

  • Cyrtomium aequibasis (C.Chr.) Ching
  • Cyrtomium anomophyllum (Zenker) Fraser-Jenk.
  • Cyrtomium atropunctatum Kurata
  • Cyrtomium caryotideum (Wall. ex Hook. & Grev.) C.Presl
  • Cyrtomium chingianum P.S.Wang
  • Cyrtomium confertifolium Ching & K.H.Shing
  • Cyrtomium conforme Ching
  • Cyrtomium devexiscapulae (Koidz.) Ching
  • Cyrtomium elongatum S.K.Wu & P.K.Lôc
  • Cyrtomium falcatum (L.f.) C.Presl (Japanese holly fern or holly fern) - type species
  • Cyrtomium fortunei J.Sm.
  • Cyrtomium grossum Christ
  • Cyrtomium guizhouense H.S.Kung & P.S.Wang
  • Cyrtomium hemionitis Christ
  • Cyrtomium laetevirens (Hiyama) Nakaike
  • Cyrtomium latifalcatum S.K.Wu & Mitsuta
  • Cyrtomium lonchitoides (Christ) Christ
  • Cyrtomium luctuosum J.P.Roux
  • Cyrtomium macrophyllum (Makino) Tagawa
  • Cyrtomium membranifolium Ching & K.H.Shing ex H.S.Kung & P.S.Wang
  • Cyrtomium micropterum (Kunze) Ching
  • Cyrtomium nephrolepioides (Christ) Copel.
  • Cyrtomium obliquum Ching & K.H.Shing
  • Cyrtomium omeiense China & Shing
  • Cyrtomium pachyphyllum (Rosenst.) C.Chr.
  • Cyrtomium pseudocaryotideum J.P.Roux
  • Cyrtomium serratum Ching & K.H.Shing
  • Cyrtomium shingianum H.S.Kung & P.S.Wang
  • Cyrtomium sinningense Ching & K.H.Shing
  • Cyrtomium taiwanianum Tagawa
  • Cyrtomium takusicola Tagawa
  • Cyrtomium tsinglingense Ching & K.H.Shing
  • Cyrtomium urophyllum Ching
  • Cyrtomium yamamotoi Tagawa
  • Cyrtomium yunnanense Ching
References

References

  1. Hassler, Michael & Schmitt, Bernd (January 2020). "Cyrtomium". Checklist of Ferns and Lycophytes of the World. 8.20. Archived from the original on 2017-09-02. Retrieved 2020-02-02.
  2. PPG I (2016). "A community-derived classification for extant lycophytes and ferns". Journal of Systematics and Evolution. 54 (6): 563–603. doi:10.1111/jse.12229. S2CID 39980610.
  3. Le Péchon, Timothée; Zhang, Liang; He, Hai; Zhou, Xin-Mao; Bytebier, Benny; Gao, Xin-Fen; Zhang, Li-Bing (May 2016). "A well-sampled phylogenetic analysis of the polystichoid ferns (Dryopteridaceae) suggests a complex biogeographical history involving both boreotropical migrations and recent transoceanic dispersals". Molecular Phylogenetics and Evolution. 98: 324–336. Bibcode:2016MolPE..98..324L. doi:10.1016/j.ympev.2016.02.018. PMID 26944012.
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