Article · Wikipedia archive · Last revised Aug 7, 2026

Chloridoideae

Chloridoideae is one of the largest subfamilies of grasses, with roughly 150 genera and 1,600 species, mainly found in arid tropical or subtropical grasslands. Within the PACMAD clade, their sister group is the Danthonioideae. The subfamily includes widespread weeds such as Bermuda grass, goosegrass or finger grass (Chloris), but also millet species grown in some tropical regions, namely finger millet and teff.

Last revised
Aug 7, 2026
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Chloridoideae
Finger millet (Eleusine coracana) in Nepal
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Embryophytes
Clade: Tracheophytes
Clade: Spermatophytes
Clade: Angiosperms
Clade: Monocots
Clade: Commelinids
Order: Poales
Family: Poaceae
Clade: PACMAD clade
Subfamily: Chloridoideae
Kunth ex Beilschm.
Tribes

Chloridoideae is one of the largest subfamilies of grasses, with roughly 150 genera and 1,600 species, mainly found in arid tropical or subtropical grasslands. Within the PACMAD clade, their sister group is the Danthonioideae.1 The subfamily includes widespread weeds such as Bermuda grass (Cynodon dactylon), goosegrass (Eleusine indica) or finger grass (Chloris), but also millet species grown in some tropical regions, namely finger millet (Eleusine coracana) and teff (Eragrostis tef).

With the exception of some species in Ellisochloa and Eleusine indica, most of the subfamily's species use the C4 photosynthetic pathway. The first evolutionary transition from C3 to C4 photosynthesis in the grasses probably occurred in this subfamily, around 32 to 25 million years ago in the Oligocene.2

Phylogeny

Relationships of tribes in the Chloridoideae according to a 2017 phylogenetic classification, also showing the Danthonioideae as sister group:3

The following genera have not been assigned to a tribe:3

References

References

  1. Soreng, Robert J.; Peterson, Paul M.; Romschenko, Konstantin; Davidse, Gerrit; Zuloaga, Fernando O.; Judziewicz, Emmet J.; Filgueiras, Tarciso S.; Davis, Jerrold I.; Morrone, Osvaldo (2015). "A worldwide phylogenetic classification of the Poaceae (Gramineae)". Journal of Systematics and Evolution. 53 (2): 117–137. doi:10.1111/jse.12150. hdl:11336/25248. ISSN 1674-4918. Open access icon
  2. Christin, Pascal-Antoine; Besnard, Guillaume; Samaritani, Emanuela; Duvall, Melvin R.; Hodkinson, Trevor R.; Savolainen, Vincent; Salamin, Nicolas (2008). "Oligocene CO2 Decline Promoted C4 Photosynthesis in Grasses". Current Biology. 18 (1): 37–43. doi:10.1016/j.cub.2007.11.058. hdl:2262/82791. ISSN 0960-9822. PMID 18160293. Open access icon
  3. Soreng, Robert J.; Peterson, Paul M.; Romaschenko, Konstantin; Davidse, Gerrit; Teisher, Jordan K.; Clark, Lynn G.; Barberá, Patricia; Gillespie, Lynn J.; Zuloaga, Fernando O. (2017). "A worldwide phylogenetic classification of the Poaceae (Gramineae) II: An update and a comparison of two 2015 classifications". Journal of Systematics and Evolution. 55 (4): 259–290. doi:10.1111/jse.12262. hdl:10261/240149. ISSN 1674-4918.